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A nuclear localization signal at the SAM-SAM domain interface of AIDA-1 suggests a requirement for domain uncoupling prior to nuclear import.
Kurabi, Arwa; Brener, Stacey; Mobli, Mehdi; Kwan, Jamie J; Donaldson, Logan W.
Afiliación
  • Kurabi A; Department of Biology, York University, Toronto, Ontario, Canada.
J Mol Biol ; 392(5): 1168-77, 2009 Oct 09.
Article en En | MEDLINE | ID: mdl-19666031
The neuronal scaffolding protein AIDA-1 is believed to act as a convener of signals arising at postsynaptic densities. Among the readily identifiable domains in AIDA-1, two closely juxtaposed sterile alpha motif (SAM) domains and a phosphotyrosine binding domain are located within the C-terminus of the longest splice variant and exclusively in four shorter splice variants. As a first step towards understanding the possible emergent properties arising from this assembly of ligand binding domains, we have used NMR methods to solve the first structure of a SAM domain tandem. Separated by a 15-aa linker, the two SAM domains are fused in a head-to-tail orientation that has been observed in other hetero- and homotypic SAM domain structures. The basic nuclear import signal for AIDA-1 is buried at the interface between the two SAM domains. An observed disparity between the thermal stabilities of the two SAM domains suggests a mechanism whereby the second SAM domain decouples from the first SAM domain to facilitate translocation of AIDA-1 to the nucleus.
Asunto(s)

Texto completo: 1 Colección: 01-internacional Base de datos: MEDLINE Asunto principal: Proteínas Portadoras / Núcleo Celular / Señales de Localización Nuclear Idioma: En Revista: J Mol Biol Año: 2009 Tipo del documento: Article País de afiliación: Canadá Pais de publicación: Países Bajos

Texto completo: 1 Colección: 01-internacional Base de datos: MEDLINE Asunto principal: Proteínas Portadoras / Núcleo Celular / Señales de Localización Nuclear Idioma: En Revista: J Mol Biol Año: 2009 Tipo del documento: Article País de afiliación: Canadá Pais de publicación: Países Bajos