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Alternative splicing facilitates internal ribosome entry on the ornithine decarboxylase mRNA.
Pyronnet, S; Pradayrol, L; Sonenberg, N.
Afiliación
  • Pyronnet S; INSERM U531, Institut Louis Bugnard IFR31, BP84225, Hôpital Rangueil, 31432 Toulouse Cedex 4, France. pyronnet@toulouse.inserm.fr
Cell Mol Life Sci ; 62(11): 1267-74, 2005 Jun.
Article en En | MEDLINE | ID: mdl-15905964
Ornithine decarboxylase (ODC) is the ratelimiting enzyme in the biosynthesis of polyamines, which are required for optimal cell growth and proliferation. ODC is overexpressed in many tumors and, conversely, its overexpression induces transformation. We have previously reported that ODC mRNA alternative splicing relieves the translation repression normally imposed by a long and structured 5' untranslated region (UTR), and that the ODC 5' UTR contains an internal ribosome entry site (IRES). Here we show that ODC IRES activity is enhanced following inclusion of alternative sequences generated by splicing at cryptic acceptor sites. Furthermore, the alternative ODC IRES is more sensitive to cell-cycledependent changes in the rate of translation. These findings uncover a new biological property of differentially spliced transcripts. This is the first example of alternative splicing that modulates mRNA translation through the cell cycle in a cap-independent manner.
Asunto(s)
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Colección: 01-internacional Base de datos: MEDLINE Asunto principal: Ornitina Descarboxilasa / Ribosomas / ARN Mensajero / Empalme Alternativo Límite: Humans Idioma: En Revista: Cell Mol Life Sci Asunto de la revista: BIOLOGIA MOLECULAR Año: 2005 Tipo del documento: Article País de afiliación: Francia Pais de publicación: Suiza
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Colección: 01-internacional Base de datos: MEDLINE Asunto principal: Ornitina Descarboxilasa / Ribosomas / ARN Mensajero / Empalme Alternativo Límite: Humans Idioma: En Revista: Cell Mol Life Sci Asunto de la revista: BIOLOGIA MOLECULAR Año: 2005 Tipo del documento: Article País de afiliación: Francia Pais de publicación: Suiza