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Amastin mRNA abundance in Trypanosoma cruzi is controlled by a 3'-untranslated region position-dependent cis-element and an untranslated region-binding protein.
Coughlin, B C; Teixeira, S M; Kirchhoff, L V; Donelson, J E.
Afiliación
  • Coughlin BC; Department of Biochemistry, University of Iowa, Iowa City 52242, USA.
J Biol Chem ; 275(16): 12051-60, 2000 Apr 21.
Article en En | MEDLINE | ID: mdl-10766837
The genome of Trypanosoma cruzi contains tandem arrays of alternating genes encoding amastin and tuzin. Amastin is a surface glycoprotein abundantly expressed on the intracellular mammalian amastigote form of the protozoan parasite, and tuzin is a G-like protein. We demonstrated previously that the amastin-tuzin gene cluster is polycistronically transcribed to an equal extent in all parasite life cycle stages. The steady state level of amastin mRNA, however, is 68-fold more abundant in amastigotes than in epimastigotes. Here we show that the half-life of amastin mRNA is 7 times longer in amastigotes than in epimastigotes. Linker replacement experiments demonstrate that the middle one-third of the 630-nucleotide 3'-untranslated region (UTR) is responsible for the amastin mRNA up-regulation. This positive effect is dependent on the distance of the 3'-UTR segment from the stop codon and the polyadenylation site as well as on its orientation. A protein or protein complex more abundant in amastigotes than in epimastigotes binds to this minimally defined 3'-UTR segment and may be involved in its regulatory function.
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Colección: 01-internacional Base de datos: MEDLINE Asunto principal: Trypanosoma cruzi / ARN Mensajero / Glicoproteínas de Membrana / Proteínas Protozoarias / Elementos de Facilitación Genéticos / Regiones no Traducidas 3' Límite: Animals Idioma: En Revista: J Biol Chem Año: 2000 Tipo del documento: Article País de afiliación: Estados Unidos Pais de publicación: Estados Unidos
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Colección: 01-internacional Base de datos: MEDLINE Asunto principal: Trypanosoma cruzi / ARN Mensajero / Glicoproteínas de Membrana / Proteínas Protozoarias / Elementos de Facilitación Genéticos / Regiones no Traducidas 3' Límite: Animals Idioma: En Revista: J Biol Chem Año: 2000 Tipo del documento: Article País de afiliación: Estados Unidos Pais de publicación: Estados Unidos