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The study was conducted within a well-managed beef cattle operation in northeastern Mexico. Each nest was weighed and dissected to obtain the plant and animal material used to build the nests. The number of materials present per nest and relative frequency were determined. Twenty-one building materials were used. Over the years, Muhlenbergia torreyi represented 85.5% of the total biomass of the nests, and Aristida longiseta, Bouteloua gracilis, Brickellia canescens, Purshia mexicana and Cirsium ehrenbergii constituted 2.45, 2.80, 2.44, 1.34 and 1.11% of the total biomass, respectively. The above-mentioned grasses represented 95.62% of the total biomass. Material of animal origin was horse and cow hair, which represented 0.84 and 0.58% of the total biomass, respectively. It was concluded that, at the study site, Muhlenbergia torreyi was a key nest-building material for the Worthen sparrow nest.

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Goat milk composition is affected by feeding, and in semiarid rangeland, information on Creole goat milk physicochemical composition is lacking. For the fulfillment of this objective, three agroecological regions (AR) considering altitude (lowland 87, highland 779, and mountain 1309 m above sea level) with different botanical compositions were chosen. Every AR analyzed accounted for 30 goat herds, with a total of 90 herds. The results demonstrated that altitude had an influence mainly on density and protein. Milk density increases as altitude increases; conversely, milk protein increases as altitude decreases. On the other hand, in the mountain and lowland ARs, the salts and solids not fat (SNF) percentages were higher compared to that of the highland AR (p < 0.05). The freezing point (FP) was higher at highland altitudes compared to that of mountain and lowland ARs (p < 0.01). In the milk fatty acids (FA) profile, only the C14:1 value was affected by altitude, whereas goat milk at lowland and mountain altitudes had higher values compared to that at highland altitudes (p < 0.05). Additionally, late lactation stage fat, FP, and pH values were higher compared to early lactation values. The opposite effect was observed for salts and SNF. In the FA profile, late lactation values were higher for C10:0 and C8:0 compared to early lactation values. The opposite trend was observed for C18:2n6t. The thrombogenic index was significantly higher at lowland altitudes compared to highland altitudes, and similar to the mountain AR. These goat milk characteristics could be explained as a consequence of animal nutrition, as well as the goat's meat-type phenotype.

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For a simple, non-invasive evaluation of nutritional status of sheep kept under extensive grazing conditions on annual rangeland, fecal indices (2,6 diaminopimelic acid, nitrogen, and phosphorus) obtained during the vegetative, reproductive, and dry grassland phenological stages, were correlated with ruminal physiological biomarkers (volatile fatty acids and ruminal ammonia). Through correlation analysis and linear regressions, the degree of association between the variables studied was established. The fecal indices that presented the highest degree of association with ruminal variables were FN and FP, being highly correlated with the production of branched-chain volatile fatty acids (isobutyrate and isovalerate) and with ruminal ammonia (r ≥ 0.65), establishing simple linear regression equations of high significance (p ≤ 0.05). Therefore, fecal indices, especially fecal concentrations of N and P, could reflect the metabolism at the ruminal level and with it the availability of compounds for microbial growth, which would help to establish the nutritional status of sheep herds under extensive grazing conditions.

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A diverse group of rangeland-medicinal plants are being used by ruminant whilst some of them have not been assessed for their nutritional value. This study was aimed to evaluate the chemical and mineral composition, buffering capacity, and in vitro fermentation of some rangeland-medicinal plants including Thymus kotschyanus, Ziziphora persica, Lallemantia royleana, and Scutellaria litwinowii in the family Lamiaceae, and Hypericum scabrum, in the family Hypericaceae. The results indicated that crude protein (CP) content ranged from 8.66% (S. litwinowii) to 12.17% of DM (H. scabrum). It was found that Z. persica had the highest potential gas production, metabolism energy (ME), relative feed value (RFV), and dry matter digestibility (DMD) values of 53.44 (mL 200-1 mg DM), 5.84 (MJ kg-1 DM), 170.66 and 70.88%, respectively. Mineral content differed among plants; Ca ranged from 5.79 to 41.96 g kg-1 DM. The concentrations of Ca, K, Mg, Fe, Zn, and Co were highest for L. royleana. Total volatile fatty acids (TVFA) and propionate concentrations were highest in the culture medium cultured with Z. persica, however, acetate, and butyrate were highest in H. scabrum. Acid-base buffering capacity was lower in T. kotschyanus and H. scabrum compared to other plants, while it was higher in S. litwinowii. Overall, it can be concluded that among plants evaluated in this study, Z. persica had higher nutritional value for sheep feeding.(AU)

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The data set covers a 101-yr period (1915-2016) of quadrat-based plant sampling at the Jornada Experimental Range in southern New Mexico. At each sampling event, a pantograph was used to record the location and perimeter of living plants within permanent quadrats. Basal area was recorded for perennial grass species, canopy cover area was recorded for shrub species, and all other perennial species were recorded as point data. The data set includes 122 1 × 1 m permanent quadrats, although not all quadrats were sampled in each year of the study and there is a gap in monitoring from 1980 to 1995. These data provide a unique opportunity to investigate changes in the plant community over 100 yr of variation in precipitation and other environmental conditions. We provide the following data and data formats: (1) the digitized maps in shapefile format; (2) a data table containing coordinates (x, y) of perennial species within quadrats, including cover area for grasses and shrubs; (3) a data table of counts of annual plant individuals per quadrat; (4) a species list indicating growth form and habit of recorded species; (5) a table of dates when each quadrat was sampled; (6) a table of the pasture each quadrat was located within (note that pasture boundaries have changed over time); (7) a table of depth to petrocalcic layer measurements taken at quadrat locations; (8) a table of particle size analysis of soil samples taken at quadrat locations; (9) a table of topographic characteristics of quadrat locations (e.g., concave or convex topography). Pantograph sampling is currently conducted at 5-yr intervals by USDA-ARS staff, and new data will be added periodically to the EDI Data Portal Repository (see section V.E.2). This information is released under the Creative Commons license-Attribution-CC BY and the consumer of these data is required to cite it appropriately in any publication that results from its use.

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C4 grasses are common species in rangelands around the world and represent an attractive option for second-generation biofuel production. Although they display high polysaccharide content and reach great levels of biomass accumulation, there is a major technical issue to be addressed before they can be used for bioethanol industrial production: lignin removal. Concerning this, Pycnoporus and Ganoderma fungal genera have been highlighted due to their ability to hydrolyze lignocellulose in biological pretreatments. Our goals here were to evaluate the pretreatment efficiency using the secretome of species from Pycnoporus and Ganoderma spp. harvested from a glucose-free inductive medium (using a C4 grass) and to identify the fungal enzymatic activities responsible for the lignin degradation and glucose release. Our results show that P. sanguineus secretome exhibits a higher activity of lignocellulolytic enzymes such as cellulases, xylanases, laccases, and manganese peroxidases compared with that from G. resinaceum. Interestingly, zymograms in the presence of 2 M glucose suggest that a ß-glucosidase isoform from P. sanguineus could be glucose tolerant. The proteomic approach carried out allowed the identification of 73 and 180 different proteins in G. resinaceum and P. sanguineus secretomes, respectively, which were functionally classified in five main categories and a miscellaneous group. These results open new avenues for future experimental work that lead to a deeper comprehension and a greater application of the mechanisms underlying lignocellulosic biomass degradation.

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The objective was to characterize milk yield and composition from mixed-breed goats kidding in winter and grazing rangelands with 221 (n = 15), 334 (n = 20), or 441 (n = 12) mm of annual rainfall in northern Mexico (22-25°N). Weekly milk yield and composition were recorded and progeny growth performance assessed. Body weight loss during lactation was the highest (6.4 kg) for goats on rangeland with 441 mm of rainfall than goats on 221 mm (1.6 kg), or 334 mm (1.8 kg; P < 0.01). The highest daily milk yield was for goats on 221 mm (498 mL; P < 0.01). There was not an evident peak of lactation and milk yield increased uninterruptedly at the end of lactation. Milk fat was negatively related to milk yield and goats on rangeland with 334 mm had the greatest milk fat content (P < 0.001). Milk protein and lactose content were higher in goats on rangeland with 221 mm (P < 0.01). Goat kids on rangelands with 221 and 441 mm grew faster and were heavier (P < 0.001) at weaning than kids reared in other rangelands. Weaning weight was positively related to milk yield (P < 0.001); but not milk components (P > 0.05). In conclusion, moderate milk yield of mixed-breed goats on rangeland is feasible with parturitions in winter, and milk yield is not affected by reduced annual precipitation in rangelands of northern Mexico. Offspring growth performance depends on maternal milk yield, but not milk composition. Nevertheless, slow growth to weaning of offspring had adverse effects on time to reach adequate market weight.

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Arbuscular mycorrhizal (AM) fungi are well-known contributors to soil aggregation and nutrient cycling functions, but we still know little about their capacity to resist or recover from persistent disturbance. Rangeland management may deteriorate these functions by affecting the activity of soil biota, including AM fungi, among other consequences. If affected, some soil properties show recovery when management stops and natural regeneration is allowed. We conducted an experiment to evaluate if the functions related to soil aggregation and promotion of exocellular enzymatic activities associated with AM fungal communities had been affected by rangeland management and, if they had, whether they recovered with successional time when management stopped. AM fungal communities from ten sites with different successional ages in a tropical dry forest region were inoculated to the same host growing in pots divided by mesh into a plant compartment and an AM mycelium compartment. We examined soil stable aggregates fractions and enzymatic activities produced or promoted by AM fungi. Soil aggregation changed significantly only after the study had run for 3 years, was higher in the hyphosphere than in the root compartment, and showed a low but positive relation with the successional age of the communities. The activity of phosphatase, but not casein-protease and beta-glucosidase, increased with successional age. Therefore, soil aggregation and enzyme activities associated with AM fungal communities seemed resilient because casein-protease and beta-glucosidase were unchanged, and aggregation and phosphatase were reduced by rangeland management but recovered with successional time.

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Ecological modeling that includes plant population processes as a critical determinant of vegetation dynamics is useful for sustainable rangeland management. However, we know little about how long-term sheep grazing pressure drives the plant community structure through changes in different native grass species at both individual and population levels. In this study, we hypothesized that plant populations perform differently under different grazing management due to their specified preference by livestock animals. We also tested whether grazing-rest management, aimed at increasing long-term rangeland sustainability, improves the plant growth of forage grass species. We evaluated plant density, individual morphology and plant-size distribution of dominant grass populations in permanent exclosures and open fields under moderate and intensive grazing pressures in Patagonian steppes (South America). We also examined the effects of seasonal grazing-rest managements on the growth and tillering (asexual reproduction) of forage species plants, using temporary mobile exclosures. Grazing intensity changed population density and structure according to species. Compared to permanent exclosures, moderate grazing maintained the plant density of palatable species highly preferred by sheep, reduced the standing-dead biomass proportion of individual plants, and promoted the green biomass of tussocks. Conversely, intensive grazing (double stocking rates) decreased the plant density and individual size of species highly preferred by sheep, and increased the plant density of non-preferred species. Grazing-rest enhanced forage grass species growth and reproduction compared with year-round grazing management, especially during the growing season of a wet year. Our studies support that sheep can be managed to control the plant-size distribution of dominant grass species, their population dynamics, and thereby the overall forage availability at the community level. Both moderate grazing and grazing-rest management can improve the forage availability and preserve the dominant native grasses. We suggest applying a plant population dynamics perspective to facilitate sustainable management of global rangelands.

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BACKGROUND: Precision Livestock Farming (PLF) is a promising approach to minimize the conflicts between socio-economic activities and landscape conservation. However, its application on extensive systems of livestock production can be challenging. The main difficulties arise because animals graze on large natural pastures where they are exposed to competition with wild herbivores for heterogeneous and scarce resources, predation risk, adverse weather, and complex topography. Considering that the 91% of the world's surface devoted to livestock production is composed of extensive systems (i.e., rangelands), our general aim was to develop a PLF methodology that quantifies: (i) detailed behavioural patterns, (ii) feeding rate, and (iii) costs associated with different behaviours and landscape traits. METHODS: For this, we used Merino sheep in Patagonian rangelands as a case study. We combined data from an animal-attached multi-sensor tag (tri-axial acceleration, tri-axial magnetometry, temperature sensor and Global Positioning System) with landscape layers from a Geographical Information System to acquire data. Then, we used high accuracy decision trees, dead reckoning methods and spatial data processing techniques to show how this combination of tools could be used to assess energy balance, predation risk and competition experienced by livestock through time and space. RESULTS: The combination of methods proposed here are a useful tool to assess livestock behaviour and the different factors that influence extensive livestock production, such as topography, environmental temperature, predation risk and competition for heterogeneous resources. We were able to quantify feeding rate continuously through time and space with high accuracy and show how it could be used to estimate animal production and the intensity of grazing on the landscape. We also assessed the effects of resource heterogeneity (inferred through search times), and the potential costs associated with predation risk, competition, thermoregulation and movement on complex topography. DISCUSSION: The quantification of feeding rate and behavioural costs provided by our approach could be used to estimate energy balance and to predict individual growth, survival and reproduction. Finally, we discussed how the information provided by this combination of methods can be used to develop wildlife-friendly strategies that also maximize animal welfare, quality and environmental sustainability.

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Grazing intensity elicits changes in the composition of plant functional groups in both shortgrass steppe (SGS) and northern mixed-grass prairie (NMP) in North America. How these grazing intensity-induced changes control aboveground net primary production (ANPP) responses to precipitation remains a central open question, especially in light of predicted climate changes. Here, we evaluated effects of four levels (none, light, moderate, and heavy) of long-term (>30 yr) grazing intensity in SGS and NMP on: (1) ANPP; (2) precipitation-use efficiency (PUE, ANPP : precipitation); and (3) precipitation marginal response (PMR; slope of a linear regression model between ANPP and precipitation). We advance prior work by examining: (1) the consequences of a range of grazing intensities (more grazed vs. ungrazed); and (2) how grazing-induced changes in ANPP and PUE are related both to shifts in functional group composition and physiological responses within each functional group. Spring (April-June) precipitation, the primary determinant of ANPP, was only 12% higher in NMP than in SGS, yet ANPP and PUE were 25% higher. Doubling grazing intensity in SGS and nearly doubling it in NMP reduced ANPP and PUE by only 24% and 33%, respectively. Increased grazing intensity reduced C3 graminoid biomass and increased C4 grass biomass in both grasslands. Functional group shifts affected PUE through biomass reductions, as PUE was positively associated with the relative abundance of C3 species and negatively with C4 species across both grasslands. At the community level, PMR was similar between grasslands and unaffected by grazing intensity. However, PMR of C3 graminoids in SGS was eightfold higher in the ungrazed treatment than under any grazed level. In NMP, PMR of C3 graminoids was only reduced under heavy grazing intensity. Knowing the ecological consequences of grazing intensity provides valuable information for mitigation and adaptation strategies in response to predicted climate change. For example, moderate grazing (the recommended rate) in SGS would sequester the same amount of aboveground carbon as light grazing because ANPP was nearly the same. In contrast, reductions in grazing intensity in NMP from moderate to light intensity would increase the amount of aboveground carbon sequestrated by 25% because of increased ANPP.

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To reduce dependence on foreign oil reserves, there has been a push in North America to develop alternative domestic energy resources. Relatively undeveloped renewable energy resources include biofuels and wind and solar energy, many of which occur predominantly on rangelands. Rangelands are also key areas for natural gas development from shales and tight sand formations. Accordingly, policies aimed at greater energy independence are likely to affect the delivery of crucial ecosystem services provided by rangelands. Assessing and dealing with the biophysical and socio-economic effects of energy development on rangeland ecosystems require an integrative and systematic approach that is predicated on a broad understanding of diverse issues related to energy development. In this article, we present a road map for developing an integrative assessment of energy development on rangelands in North America. We summarize current knowledge of socio-economic and biophysical aspects of rangeland based energy development, and we identify knowledge gaps and monitoring indicators to fill these knowledge gaps.

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A intensidade de pastejo e suas alterações ao longo do tempo podem afetar a composição botânica e a estrutura do pasto. Este trabalho buscou avaliar o efeito de distintas ofertas de forragem (OF) sobre a dinâmica populacional de espécies de uma pastagem natural dos Campos Sulinos. As avaliações da composição botânica foram realizadas em 12 de janeiro e 17 de julho através do método BOTANAL. Os tratamentos constaram de diferentes OF (kg MS 100kg-1 de peso vivo dia-1): 8, 12 e 16 durante todo ano e diferentes combinações de sequências: 8 na primavera e 12 no resto do ano (8-12); 12 na primavera e 8 no resto do ano (12-8); 16 na primavera e 12 no resto do ano (16-12). O delineamento utilizado foi o de blocos casualizados com duas repetições. A oferta de forragem provocou alterações na composição botânica da vegetação. A manipulação da OF na primavera alterou a composição botânica da vegetação e apresentou maior impacto nas menores OF.

Grazing intensity manipulation can affect structure and botanical composition of the pasture. The objective of this research was to evaluate the effects of different forage allowances and different seasonal sequences of forage allowance (FA) on the species population dynamics in the rangelands of Campos Sulinos. The botanical composition evaluations were made in January 12th and July 17th, using BOTANAL method. Treatments were of different FA (kg DM 100kg-1 of live weight day-1): 8, 12, 16 all around the year; and different sequences of FA: 8 on spring and 12 at the rest of year (8-12); 12 on spring and 8 at the rest of year (12-8) and 16 on spring and 12 at the rest of year (16-12). The statistical design used was randomized blocks with two replications. The forage allowance led to changes in botanical and structural composition of vegetation. It can be inferred that the forage allowance variation on the vegetation had higher impact on smaller forage allowances.

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Grazing intensity manipulation can affect structure and botanical composition of the pasture. The objective of this research was to evaluate the effects of different forage allowances and different seasonal sequences of forage allowance (FA) on the species population dynamics in the rangelands of Campos Sulinos. The botanical composition evaluations were made in January 12th and July 17th, using BOTANAL method. Treatments were of different FA (kg DM 100kg-1 of live weight day-1): 8, 12, 16 all around the year; and different sequences of FA: 8 on spring and 12 at the rest of year (8-12); 12 on spring and 8 at the rest of year (12-8) and 16 on spring and 12 at the rest of year (16-12). The statistical design used was randomized blocks with two replications. The forage allowance led to changes in botanical and structural composition of vegetation. It can be inferred that the forage allowance variation on the vegetation had higher impact on smaller forage allowances.

A intensidade de pastejo e suas alterações ao longo do tempo podem afetar a composição botânica e a estrutura do pasto. Este trabalho buscou avaliar o efeito de distintas ofertas de forragem (OF) sobre a dinâmica populacional de espécies de uma pastagem natural dos Campos Sulinos. As avaliações da composição botânica foram realizadas em 12 de janeiro e 17 de julho através do método BOTANAL. Os tratamentos constaram de diferentes OF (kg MS 100kg-1 de peso vivo dia-1): 8, 12 e 16 durante todo ano e diferentes combinações de sequências: 8 na primavera e 12 no resto do ano (8-12); 12 na primavera e 8 no resto do ano (12-8); 16 na primavera e 12 no resto do ano (16-12). O delineamento utilizado foi o de blocos casualizados com duas repetições. A oferta de forragem provocou alterações na composição botânica da vegetação. A manipulação da OF na primavera alterou a composição botânica da vegetação e apresentou maior impacto nas menores OF.

RESUMO

Grazing intensity manipulation can affect structure and botanical composition of the pasture. The objective of this research was to evaluate the effects of different forage allowances and different seasonal sequences of forage allowance (FA) on the species population dynamics in the rangelands of Campos Sulinos. The botanical composition evaluations were made in January 12th and July 17th, using BOTANAL method. Treatments were of different FA (kg DM 100kg-1 of live weight day-1): 8, 12, 16 all around the year; and different sequences of FA: 8 on spring and 12 at the rest of year (8-12); 12 on spring and 8 at the rest of year (12-8) and 16 on spring and 12 at the rest of year (16-12). The statistical design used was randomized blocks with two replications. The forage allowance led to changes in botanical and structural composition of vegetation. It can be inferred that the forage allowance variation on the vegetation had higher impact on smaller forage allowances.

A intensidade de pastejo e suas alterações ao longo do tempo podem afetar a composição botânica e a estrutura do pasto. Este trabalho buscou avaliar o efeito de distintas ofertas de forragem (OF) sobre a dinâmica populacional de espécies de uma pastagem natural dos Campos Sulinos. As avaliações da composição botânica foram realizadas em 12 de janeiro e 17 de julho através do método BOTANAL. Os tratamentos constaram de diferentes OF (kg MS 100kg-1 de peso vivo dia-1): 8, 12 e 16 durante todo ano e diferentes combinações de sequências: 8 na primavera e 12 no resto do ano (8-12); 12 na primavera e 8 no resto do ano (12-8); 16 na primavera e 12 no resto do ano (16-12). O delineamento utilizado foi o de blocos casualizados com duas repetições. A oferta de forragem provocou alterações na composição botânica da vegetação. A manipulação da OF na primavera alterou a composição botânica da vegetação e apresentou maior impacto nas menores OF.

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Under certain environmental conditions, several species of grasshoppers inhabiting rangeland areas (e.g., inter-valleys) of the Chilean transitional desert can irrupt demographically. One of them is Elasmoderus wagenknechti (Liebermann), an endemic species. We studied two outbreaks occurred in 1996 and 1999. The objectives of the research were to (i) estimate some demographic parameters associated with the aforementioned events, and (ii) compare between them the population parameters. The parameters we studied were density (ind/m²), sexual proportion, female fertility (i.e., eggs/female), and the relationship eggs/female versus female size. Density (ind/m²) ranged from 0.2 to 0.4 in 1996 and from 0.5 to 0.6 in 1999. ANOVA-tests registered differences in density among study sites and between years. In some of the study sites, females were more abundant than males (e.g., Lomas del Sauce). Females fertility ranged from 27 to 39 eggs per female, with significant differences among sites in 1996 but not in 1999. For females of 30-50 mm body size, a linear relationship was detected between egg number/female and body size. It is concluded that the E. wagenknechti outbreaks are site-specific in the study area and that the demographic parameters examined show a high variability among sites and between events.

Sob certas condições ambientais, diversas espécies de gafanhotos que habitam áreas de campos, como as áreas entre vales do deserto transicional do Chile, podem experimentar surtos populacionais. Uma dessas espécies é Elasmoderus wagenknechti (Liebermann), espécie endêmica. Nós estudamos dois surtos que ocorreram em 1996 e 1999. Os objetivos da pesquisa foram (1) estimar alguns parâmetros demográficos associados aos eventos acima indicados, e (2) comparar os parâmetros entre os eventos. Os parâmetros estudados foram densidade (indivíduos/m²), razão sexual, fertilidade da fêmea (i.e., ovos/fêmea), e a relação entre a fertilidade e o tamanho corporal da fêmea. O intervalo da densidade do evento foi estimado entre 0,2 e 0,4 indivíduos/m² em 1996, e entre 0,5 e 0,6 indivíduos /m² em 1999. Foram encontradas diferenças (análise de variância) na densidade entre os locais para o mesmo evento, e entre os eventos. As fêmeas foram mais numerosas que os machos em alguns locais estudados (e.g., Lomas del Sauce). Foi estimado um intervalo médio de 27-39 ovos/fêmea, existindo diferenças entre os locais no evento de 1996, mas não no evento de 1999. Para fêmeas de longitude corporal de 30-35 mm. foi detectada uma relação linear entre número de ovos/fêmea e tamanho corporal. Em conclusão, a magnitude do evento na área de estudo é específica do local; além do que para os parâmetros demográficos examinados existe uma acentuada variabilidade entre locais e entre os eventos.

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Seed size is a crucial plant trait that may potentially affect not only immediate seedling success but also the subsequent generation. We examined variation in seed weight of Wyoming sagebrush (Artemisia tridentata ssp. wyomingensis Beetle and Young), an excellent candidate species for rangeland restoration. The working hypothesis was that a major fraction of spatial and temporal variability in seed size (weight) of Wyoming sagebrush could be explained by variations in mean monthly temperatures and precipitation. Seed collection was conducted at Battle Mountain and Eden Valley sites in northern Nevada, USA, during November of 2002 and 2003. Frequency distributions of seed weight varied from leptokurtic to platykurtic, and from symmetry to skewness to the right for both sites and years. Mean seed weight varied by a factor of 1.4 between locations and years. Mean seed weight was greater (P<0.05) in 2003 than in 2002 at both sites. This can partially be attributed to 55% greater precipitation in 2003 than 2002, since mean monthly temperatures were similar (P>0.05) in all study situations. Simple linear regression showed that monthly precipitation (March to November) explained 85% of the total variation in mean seed weight ( P=0.079). Since the relationship between mean monthly temperature (June-November) and mean seed weight was not significant (r²=0.00, P=0.431), this emphasizes the importance of precipitation as an important determinant of mean seed weight. Our results suggest that the precipitation regime to which the mother plant is exposed can have a significant effect on sizes of seeds produced. Hence, seasonal changes in water availability would tend to alter size distributions of produced offspring.

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