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Agents usually adjust their strategic behaviors based on their own payoff and aspiration in gaming environments. Hence, aspiration-based learning rules play an important role in the evolutionary dynamics in a population of competing agents. However, there exist different options for how to use the aspiration information for specifying the microscopic learning rules. It is also interesting to investigate under what conditions the aspiration-based learning rules can favor the emergence of cooperative behavior in population games. A new learning rule, called as "Satisfied-Cooperate, Unsatisfied-Defect", is proposed here, which is based on aspiration. Under this learning rule, agents prefer to cooperate when their income is satisfied; otherwise, they prefer the strategy of defection. We introduce this learning rule to a population of agents playing a generalized two-person game. We, respectively, obtain the mathematical conditions in which cooperation is more abundant in finite well-mixed, infinite well-mixed, and structured populations under weak selection. Interestingly, we find that these conditions are identical, no matter whether the aspiration levels for cooperators and defectors are the same or not. Furthermore, we consider the prisoner's dilemma game (PDG) as an example and perform numerical calculations and computer simulations. Our numerical and simulation results agree well and both support our theoretical predictions in the three different types of populations. We further find that our aspiration-based learning rule can promote cooperation more effectively than alternative aspiration-based learning rules in the PDG.

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Multiplayer games on graphs are at the heart of theoretical descriptions of key evolutionary processes that govern vital social and natural systems. However, a comprehensive theoretical framework for solving multiplayer games with an arbitrary number of strategies on graphs is still missing. Here, we solve this by drawing an analogy with the Balls-and-Boxes problem, based on which we show that the local configuration of multiplayer games on graphs is equivalent to distributing k identical co-players among n distinct strategies. We use this to derive the replicator equation for any n-strategy multiplayer game under weak selection, which can be solved in polynomial time. As an example, we revisit the second-order free-riding problem, where costly punishment cannot truly resolve social dilemmas in a well-mixed population. Yet, in structured populations, we derive an accurate threshold for the punishment strength, beyond which punishment can either lead to the extinction of defection or transform the system into a rock-paper-scissors-like cycle. The analytical solution also qualitatively agrees with the phase diagrams that were previously obtained for non-marginal selection strengths. Our framework thus allows an exploration of any multi-strategy multiplayer game on regular graphs.

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The public goods game is a broadly used paradigm for studying the evolution of cooperation in structured populations. According to the basic assumption, the interaction graph determines the connections of a player where the focal actor forms a common venture with the nearest neighbors. In reality, however, not all of our partners are involved in every game. To elaborate this observation, we propose a model where individuals choose just some selected neighbors from the complete set to form a group for public goods. We explore the potential consequences by using a pair-approximation approach in a weak selection limit. We theoretically analyze how the number of total neighbors and the actual size of the restricted group influence the critical enhancement factor where cooperation becomes dominant over defection. Furthermore, we systematically compare our model with the traditional setup and show that the critical enhancement factor is lower than in the case when all players are present in the social dilemma. Hence, the suggested restricted interaction mode offers a better condition for the evolution of cooperation. Our theoretical findings are supported by numerical calculations.

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While actors in a population can interact with anyone else freely, social relations significantly influence our inclination toward particular individuals. The consequence of such interactions, however, may also form the intensity of our relations established earlier. These dynamical processes are captured via a coevolutionary model staged in multiplex networks with two distinct layers. In a so-called relationship layer, the weights of edges among players may change in time as a consequence of games played in the alternative interaction layer. As an reasonable assumption, bilateral cooperation confirms while mutual defection weakens these weight factors. Importantly, the fitness of a player, which basically determines the success of a strategy imitation, depends not only on the payoff collected from interactions, but also on the individual relationship index calculated from the mentioned weight factors of related edges. Within the framework of weak prisoner's dilemma situation, we explore the potential outcomes of the mentioned coevolutionary process where we assume different topologies for relationship layer. We find that higher average degree of the relationship graph is more beneficial to maintain cooperation in regular graphs, but the randomness of links could be a decisive factor in harsh situations. Surprisingly, a stronger coupling between relationship index and fitness discourage the evolution of cooperation by weakening the direct consequence of a strategy change. To complete our study, we also monitor how the distribution of relationship index vary and detect a strong relation between its polarization and the general cooperation level.

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Competing strategies in an evolutionary game model, or species in a biosystem, can easily form a larger unit which protects them from the invasion of an external actor. Such a defensive alliance may have two, three, four or even more members. But how effective can be such formation against an alternative group composed by other competitors? To address this question we study a minimal model where a two-member and a four-member alliances fight in a symmetric and balanced way. By presenting representative phase diagrams, we systematically explore the whole parameter range which characterizes the inner dynamics of the alliances and the intensity of their interactions. The group formed by a pair, who can exchange their neighboring positions, prevail in the majority of the parameter region. The rival quartet can only win if their inner cyclic invasion rate is significant while the mixing rate of the pair is extremely low. At specific parameter values, when neither of the alliances is strong enough, new four-member solutions emerge where a rock-paper-scissors-like trio is extended by the other member of the pair. These new solutions coexist hence all six competitors can survive. The evolutionary process is accompanied by serious finite-size effects which can be mitigated by appropriately chosen prepared initial states.

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Human society and natural environment form a complex giant ecosystem, where human activities not only lead to the change in environmental states, but also react to them. By using collective-risk social dilemma game, some studies have already revealed that individual contributions and the risk of future losses are inextricably linked. These works, however, often use an idealistic assumption that the risk is constant and not affected by individual behaviors. Here, we develop a coevolutionary game approach that captures the coupled dynamics of cooperation and risk. In particular, the level of contributions in a population affects the state of risk, while the risk in turn influences individuals' behavioral decision-making. Importantly, we explore two representative feedback forms describing the possible effect of strategy on risk, namely, linear and exponential feedbacks. We find that cooperation can be maintained in the population by keeping at a certain fraction or forming an evolutionary oscillation with risk, independently of the feedback type. However, such evolutionary outcome depends on the initial state. Taken together, a two-way coupling between collective actions and risk is essential to avoid the tragedy of the commons. More importantly, a critical starting portion of cooperators and risk level is what we really need for guiding the evolution toward a desired direction.

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According to the evolutionary death-birth protocol, a player is chosen randomly to die and neighbors compete for the available position proportional to their fitness. Hence, the status of the focal player is completely ignored and has no impact on the strategy update. In this paper, we revisit and generalize this rule by introducing a weight factor to compare the payoff values of the focal and invading neighbors. By means of evolutionary graph theory, we analyze the model on joint transitive graphs to explore the possible consequences of the presence of a weight factor. We find that focal weight always hinders cooperation under weak selection strength. Surprisingly, the results show a nontrivial tipping point of the weight factor where the threshold of cooperation success shifts from positive to negative infinity. Once focal weight exceeds this tipping point, cooperation becomes unreachable. Our theoretical predictions are confirmed by Monte Carlo simulations on a square lattice of different sizes. We also verify the robustness of the conclusions to arbitrary two-player prisoner's dilemmas, to dispersal graphs with arbitrary edge weights, and to interaction and dispersal graphs overlapping arbitrarily.

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The application of incentives, such as reward and punishment, is a frequently applied way for promoting cooperation among interacting individuals in structured populations. However, how to properly use the incentives is still a challenging problem for incentive-providing institutions. In particular, since the implementation of incentive is costly, to explore the optimal incentive protocol, which ensures the desired collective goal at a minimal cost, is worthy of study. In this work, we consider the positive and negative incentives for a structured population of individuals whose conflicting interactions are characterized by a Prisoner's Dilemma game. We establish an index function for quantifying the cumulative cost during the process of incentive implementation, and theoretically derive the optimal positive and negative incentive protocols for cooperation on regular networks. We find that both types of optimal incentive protocols are identical and time-invariant. Moreover, we compare the optimal rewarding and punishing schemes concerning implementation cost and provide a rigorous basis for the usage of incentives in the game-theoretical framework. We further perform computer simulations to support our theoretical results and explore their robustness for different types of population structures, including regular, random, small-world and scale-free networks.

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Motion is a typical reaction among animals and humans trying to reach better conditions in a changing world. This aspect has been studied intensively in social dilemmas where competing players' individual and collective interests are in conflict. Starting from the traditional public goods game model, where players are locally fixed and unconditional cooperators or defectors are present, we introduce two additional strategies through which agents can change their positions of dependence on the local cooperation level. More importantly, these so-called sophisticated players should bear an extra cost to maintain their permanent capacity to evaluate their neighborhood and react accordingly. Hence, four strategies compete, and the most successful one can be imitated by its neighbors. Crucially, the introduction of costly movement has a highly biased consequence on the competing main strategies. In the majority of parameter space, it is harmful to defectors and provides a significantly higher cooperation level when the population is rare. At an intermediate population density, which would be otherwise optimal for a system of immobile players, the presence of mobile actors could be detrimental if the interaction pattern changes slightly, thereby blocking the optimal percolation of information flow. In this parameter space, sophisticated cooperators can also show the co-called Moor effect by first avoiding the harmful vicinity of defectors; they subsequently transform into an immobile cooperator state. Hence, paradoxically, the additional cost of movement could be advantageous to reach a higher general income, especially for a rare population when subgroups would be isolated otherwise.

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Opinion dynamics on social networks have received considerable attentions in recent years. Nevertheless, just a few works have theoretically analyzed the condition in which a certain opinion can spread in the whole structured population. In this article, we propose an evolutionary game approach for a binary opinion model to explore the conditions for an opinion's spreading. Inspired by real-life observations, we assume that an agent's choice to select an opinion is not random but is based on a score rooted from both public knowledge and the interactions with neighbors. By means of coalescing random walks, we obtain a condition in which opinion A can be favored to spread on social networks in the weak selection limit. We find that the successfully spreading condition of opinion A is closely related to the basic scores of binary opinions, the feedback scores on opinion interactions, and the structural parameters including the edge weights, the weighted degrees of vertices, and the average degree of the network. In particular, when individuals adjust their opinions based solely on the public information, the vitality of opinion A depends exclusively on the difference of basic scores of A and B. When there are no negative (positive) feedback interactions between connected individuals, we find that the success of opinion A depends on the ratio of the obtained positive (negative) feedback scores of competing opinions. To complete our study, we perform computer simulations on fully connected, small-world, and scale-free networks, respectively, which support and confirm our theoretical findings.

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Explaining the emergence and maintenance of cooperation among selfish individuals from an evolutionary perspective remains a grand challenge in biology, economy and social sciences. Social exclusion is believed to be an answer to this conundrum. However, previously related studies often assume one-shot interactions and ignore how free-riding is identified, which seem to be too idealistic. In this work, we consider repeated interactions where excluders need to pay a monitoring cost to identify free-riders for exclusion and free-riders cannot participate in the following possible game interactions once they are identified and excluded by excluders in the repeated interaction process. We reveal that the introduction of such exclusion can prevent the breakdown of cooperation in repeated group interactions. In particular, we demonstrate that an evolutionary oscillation among cooperators, defectors and excluders can appear in infinitely large populations when early exclusion is implemented. In addition, we find that the population spends most of the time in states where cooperators dominate for early exclusion when stochastic mutation-selection is considered in finite populations. Our results highlight that early exclusion is successful in solving the mentioned enigma of cooperation in repeated group interactions.

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Finding appropriate incentives to enforce collaborative efforts for governing the commons in risky situations is a long-lasting challenge. Previous works have demonstrated that both punishing free-riders and rewarding cooperators could be potential tools to reach this goal. Despite weak theoretical foundations, policy makers frequently impose a punishment-reward combination. Here, we consider the emergence of positive and negative incentives and analyze their simultaneous impact on sustaining risky commons. Importantly, we consider institutions with fixed and flexible incentives. We find that a local sanctioning scheme with pure reward is the optimal incentive strategy. It can drive the entire population toward a highly cooperative state in a broad range of parameters, independently of the type of institutions. We show that our finding is also valid for flexible incentives in the global sanctioning scheme, although the local arrangement works more effectively.

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The conflict between individual and collective interests is in the heart of every social dilemmas established by evolutionary game theory. We cannot avoid these conflicts but sometimes we may choose which interaction framework to use as a battlefield. For instance some people like to be part of a larger group while other persons prefer to interact in a more personalized, individual way. Both attitudes can be formulated via appropriately chosen traditional games. In particular, the prisoner's dilemma game is based on pair interaction while the public goods game represents multi-point interactions of group members. To reveal the possible advantage of a certain attitude we extend these models by allowing players not simply to change their strategies but also let them to vary their attitudes for a higher individual income. We show that both attitudes could be the winner at a specific parameter value. Interestingly, however, the subtle interplay between different states may result in a counterintuitive evolutionary outcome where the increase of the multiplication factor of public goods game drives the population to a fully defector state. We point out that the accompanying pattern formation can only be understood via the multipoint or multi-player interactions of different microscopic states where the vicinity of a particular state may influence the relation of two other competitors.

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Varying environmental conditions affect relations between interacting individuals in social dilemmas, thus affecting also the evolution of cooperation. Oftentimes these environmental variations are seasonal and can therefore be mathematically described as periodic changes. Accordingly, we here study how periodic shifts between different manifestations of social dilemmas affect cooperation. We observe a non-trivial interplay between the inherent spatiotemporal dynamics that characterizes the spreading of cooperation in a particular social dilemma type and the frequency of payoff changes. In particular, we show that periodic changes between two available games with global ordering best be fast, while periodic changes between global and local ordering games best be slow for cooperation to thrive. We also show that the frequency of periodic changes between two local ordering social dilemmas is irrelevant, because then the process is fast and simply the average cooperation level of the two is returned. The structure of the interaction network plays an important role too in that lattices promote local ordering, whilst random graphs hinder the formation of compact cooperative clusters. Conversely, for local ordering the regular structure of the interaction network is only marginally relevant as role-separating checkerboard patterns do not rely on long-range order.

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Cooperation is the cornerstone of human evolutionary success. Like no other species, we champion the sacrifice of personal benefits for the common good, and we work together to achieve what we are unable to achieve alone. Knowledge and information from past generations is thereby often instrumental in ensuring we keep cooperating rather than deteriorating to less productive ways of coexistence. Here we present a mathematical model based on evolutionary game theory that shows how using the past as the benchmark for evolutionary success, rather than just current performance, significantly improves cooperation in the future. Interestingly, the details of just how the past is taken into account play only second-order importance, whether it be a weighted average of past payoffs or just a single payoff value from the past. Cooperation is promoted because information from the past disables fast invasions of defectors, thus enhancing the long-term benefits of cooperative behavior.

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According to the evolutionary game theory principle, a strategy representing a higher payoff can spread among competitors. But there are cases when a player consistently overestimates or underestimates her own payoff, which undermines proper comparison. Interestingly, both underconfident and overconfident individuals are capable of elevating the cooperation level significantly. While former players stimulate a local coordination of strategies, the presence of overconfident individuals enhances the spatial reciprocity mechanism. In both cases the propagations of competing strategies are influenced in a biased way resulting in a cooperation supporting environment. These effects are strongly related to the nonlinear character of invasion probabilities which is a common and frequently observed feature of microscopic dynamics.

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Utilizing common resources is always a dilemma for community members. While cooperator players restrain themselves and consider the proper state of resources, defectors demand more than their supposed share for a higher payoff. To avoid the tragedy of the common state, punishing the latter group seems to be an adequate reaction. This conclusion, however, is less straightforward when we acknowledge the fact that resources are finite and even a renewable resource has limited growing capacity. To clarify the possible consequences, we consider a coevolutionary model where beside the payoff-driven competition of cooperator and defector players the level of a renewable resource depends sensitively on the fraction of cooperators and the total consumption of all players. The applied feedback-evolving game reveals that beside a delicately adjusted punishment it is also fundamental that cooperators should pay special attention to the growing capacity of renewable resources. Otherwise, even the usage of tough punishment cannot save the community from an undesired end.

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Detecting defection and alarming partners about the possible danger could be essential to avoid being exploited. This act, however, may require a huge individual effort from those who take this job, hence such a strategy seems to be unfavorable. But structured populations can provide an opportunity where a largely unselfish excluder strategy can form an effective alliance with other cooperative strategies, hence they can sweep out defection. Interestingly, this alliance is functioning even at the extremely high cost of exclusion where the sole application of an exclusion strategy would be harmful otherwise. These results may explain why the emergence of extreme selfless behavior is not necessarily against individual selection but could be the result of an evolutionary process.

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Prosocial punishment has been proved to be a powerful mean to promote cooperation. Recent studies have found that social exclusion, which indeed can be regarded as a kind of punishment, can also support cooperation. However, if prosocial punishment and exclusion are both present, it is still unclear which strategy is more advantageous to curb free-riders. Here we first study the direct competition between different types of punishment and exclusion. We find that pool (peer) exclusion can always outperform pool (peer) punishment both in the optional and in the compulsory public goods game, no matter whether second-order sanctioning is considered or not. Furthermore, peer exclusion does better than pool exclusion both in the optional and in the compulsory game, but the situation is reversed in the presence of second-order exclusion. Finally, we extend the competition among all possible sanctioning strategies and find that peer exclusion can outperform all other strategies in the absence of second-order exclusion and punishment, while pool exclusion prevails when second-order sanctioning is possible. Our results demonstrate that exclusion is a more powerful strategy than punishment for the resolution of social dilemmas.

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''Three is a crowd" is an old proverb that applies as much to social interactions as it does to frustrated configurations in statistical physics models. Accordingly, social relations within a triangle deserve special attention. With this motivation, we explore the impact of topological frustration on the evolutionary dynamics of the snowdrift game on a triangular lattice. This topology provides an irreconcilable frustration, which prevents anticoordination of competing strategies that would be needed for an optimal outcome of the game. By using different strategy updating protocols, we observe complex spatial patterns in dependence on payoff values that are reminiscent to a honeycomb-like organization, which helps to minimize the negative consequence of the topological frustration. We relate the emergence of these patterns to the microscopic dynamics of the evolutionary process, both by means of mean-field approximations and Monte Carlo simulations. For comparison, we also consider the same evolutionary dynamics on the square lattice, where of course the topological frustration is absent. However, with the deletion of diagonal links of the triangular lattice, we can gradually bridge the gap to the square lattice. Interestingly, in this case the level of cooperation in the system is a direct indicator of the level of topological frustration, thus providing a method to determine frustration levels in an arbitrary interaction network.

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