RESUMEN
The hyporheic zone is an ecologically important ecotone that describes the extent to which nutrient-rich surface waters penetrate the shallow subsurface adjacent to a flowing surface water body. Although steady-state models satisfactorily explain the incursion of surface water into the subsurface as a function of head gradients developed across streambed riffles, they fail to account for the depth that surface water is observed to penetrate the subsurface or for the extent to which the hyporheic zone develops adjacent to the stream channel. To investigate these issues, transient flow modeling has been conducted at the riffle scale and supported by data for an instrumented site in northern Ontario where stream-stage fluctuations are strictly regulated. Model results show that daily stream-stage fluctuations between 0.6 and 4 m produce oscillating solute flow paths that typically reduce residence times of water and solutes in the hyporheic zone from 60 days or more under steady-state conditions to less than 1 day. Furthermore, similar stream-stage fluctuations increase the depth that solutes pervade the subsurface and banks lateral to the stream from around 1 m under steady-state conditions to as much as 2 and 10 m, respectively. The results demonstrate that the transient flow conditions triggered in the subsurface by variable stream stage can exert a strong influence on hyporheic zone development and have important implications for the hyporheos. The results are especially important for hyporheic communities that may survive gradual changes to their living conditions by migrating to more hospitable aquatic habitats, but are unable to respond to rapid changes provoked by more extreme hydrological events.